England broke up a rich family heritage in Tongan rugby by picking young prop Mako Vunipola on Thursday as one of four changes to its 32-man senior squad for the autumn internationals.
Loose forward James Haskell, winger Ugo Monye and hooker Tom Youngs - the brother of scrumhalf Ben - were also brought in for the matches against Fiji, Australia, South Africa and New Zealand, which are crucial to England as it looks to secure a top-four seeding ahead of the pool draw for the 2015 World Cup in December.
But it was the selection of the giant Vunipola that stood out when head coach Stuart Lancaster revealed his revised elite squad.
The Saracens forward was born in New Zealand but his father, six uncles and grandfather all played for Tonga. However, the 21-year-old Mako has pledged his allegiance to England, having already starred for the country's under-20 team, and is tipped for a huge future.
"We've watched him very closely and I'm looking for a fault in his game. Can't find one at the moment," said England forwards coach Graham Rowntree, a former prop with the national team. "I have no doubt he'll make his name in this autumn series."
Vunipola wasn't even playing professional rugby 18 months ago and spent the whole of his first year at Saracens last season in the treatment room, meaning his rise to the international stage has been meteoric. He has mixed it with the best of his prop peers in the Heineken Cup, Europe's top club competition, over the past month.
"I like his commitment to every aspect of the game; he's a dynamic ball-carrier, loves a scrum - and you know that turns me on - respects the hard yards, the tackling, the clearing out. He has his priorities right," Rowntree added. "I'm sure he'll take his chance."
Vunipola, one of four props in the squad, will start on the bench against Fiji on Nov. 10 but making an impact as a replacement could force the England coaching staff to think seriously to starting him against the three southern hemisphere powers over the following three weekends.
Tom Youngs' call-up is equally impressive, considering he has mastered the difficult role of hooker in the space of only three years after swapping from playing as a center at Leicester Tigers.
Monye is back, two years after winning his last cap, following some stand-out performances for English champion Harlequins and replaces fullback Ben Foden. Forwards Alex Corbisiero, Tom Croft and Rob Webber are also injured, allowing Lancaster to make changes to his elite squad.
The coaches have elected against picking anyone playing in France, sticking to the recently adopted policy of going down that route only under "exceptional circumstances." That means Toulon's Steffon Armitage, arguably the form English flanker, is overlooked as is his teammate Andrew Sheridan.
Lancaster was hired as full-time head coach after impressing the RFU with his inspirational qualities during last year's Six Nations, where he instilled discipline into a team that emerged from a calamitous 2011 World Cup campaign with a tarnished image after off-the-field incidents.
However, the honeymoon period is over and after losing 2-0 in South Africa in a three-match summer series, England needs results against the world's top teams this autumn not just to sustain morale but also to stay fourth in the IRB rankings above France.
That would ensure being among the top seeds for the pool draw for the 2015 World Cup - hosted by England - and avoiding the All Blacks, Wallabies and Springboks in the group stage.
"We get every week, every game, every session right and the rankings will take care of themselves," Rowntree said.
"I think we are learning, still a young group - players and coaching staff - moving forward. We are happy with where we are at the moment and what we've learnt from the summer tour, what we've learnt from each other."
Lancaster will name his captain on Tuesday, during the squad's five-day stint training at English football's new national center in Burton, central England. Flanker Chris Robshaw is expected to keep the armband.
"It will be a big test for our character and resolve, not just playing against the best in the world but to do it week in, week out," England backs coach Andy Farrell said.
ENGLAND: Forwards: Mouritz Botha, Dan Cole, James Haskell, Phil Dowson, Dylan Hartley, Tom Johnson, Courtney Lawes, Joe Marler, Ben Morgan, Tom Palmer, Geoff Parling, Chris Robshaw, Mako Vunipola, Thomas Waldrom, David Wilson, Tom Wood, Tom Youngs. Backs: Anthony Allen, Chris Ashton, Brad Barritt, Mike Brown, Danny Care, Lee Dickson, Owen Farrell, Toby Flood, Alex Goode, Jonathan Joseph, Ugo Monye, Charlie Sharples, Manu Tuilagi, Jordan Turner-Hall, Ben Youngs.